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News

 

 

JANUARY 2006

Following the last decision of the MIC Academic Consul, all Online students, once registered, can join in MIC Paris School,  in MIC School in Almaty or MIC Canada for an Onsite Program.

 

NOVEMBER 2005

The First MBA group started in November 2005 at IFCM in Paris. The second session will start in February 2006.

Students would travel to Canada for the last semester.  The program is held at MIC in Montreal or Ottawa, where students combine one semester of intensive study with an internship program within a Canadian company.

MAY 2005

The First MIC MBA Program in Paris, France, is planned to start in October 2005. The MIC registered Students from all units in the world can be transfered and study in our school in Paris. 

For more information, please contact MIC main office in Montreal.

 

FEBRUARY 2005

MIC created a school of management in Paris, France, in partnership with Group COGEFI, a well known French Business School.

FRANCO-CANADIAN INSTITUTE  (IFC) (Institut Franco-Canadien - IFC) represents this partnership.

IFC offers MBA (Master in business Administration)’ and ‘Master Européen de Management’.  The MBA program in Paris is now offered in English and in French.

To visit the MIC Paris School, please click here.

- The MBA Program in Almaty, Kazakhstan  is supposed to be started in June 2005. All Central Asian and Eastern European applicants can apply at Turan University: www.turan.kz

(http://www.turan.kz/canada/canada.html)

 

-MIC in Durban, South Africa, is offering only Online MBA Program.

 

 

NOVEMBER 2004

- MIC has signed an agreement with the Turan University in Almaty, Kazakhstan, to offer an MBA On-Campus Program in Almaty and in Astana (Capital of Kazakhstan).

Starting date: June 2005.

- Following the last decision of the Academic Consul, MIC is restructuring the teaching system for the Online EMBA and MPA programs.

The new system would start from the beginning of the New Year.

 

 

OCTOBER 2004

- Another partnership with a university in the North of Russia is the processes of negotiation. This partnership is based on offering a joint MBA On Campus Program.

Starting date is supposed to be in September  2005.                                    

More details will be published soon.

SEPTEMBER 2004

- Till September 2004, all students at MIC for the Overseas Program were from Eastern Europe and Central Asia. MIC has admitted for the first time new applicants from Saudi Arabia. The Management welcomes to the new students.

The current MIC Online Students from Overseas are from these coutries:

 

Albania

Ukraine 

Uzbekistan

Bulgaria

Kyrgyzstan

Kazakhstan

Azerbaijan

Saudi Arabia

 -MIC is finalizing an important partnership with one of the largest universities in Central Asia, Kazakhstan.

This partnership is based on offering an MBA Program On Campus Program.

Starting date is supposed to be in January 2005. More details will be published soon.

 AUGUST 2004

- MIC has developed a VIRTUAL- CLASS system. Professors from Canada will be in direct connection with students from all over the world and will teach them directly through  Internet. The V-Class system will be implemented from January 2005.

- Starting from June 2004, MIC has increased the tuition fee for eMBA and MPA programs. All students who have applied online before this date, having received a pre-admission letter, will benefit from the previous rate upon request.

 

 Montreal

 

rôle du notaire

 

 

 

World Education System

This page edited from MIT webpage

http://72.14.209.104/search?q=cache:JuUEExw0Re4J:massachusetts.college.of.liberal.arts.en.infovx.biz/+=en&gl=ca&ct=clnk&cd=106

 

Bachelor of Science

A Bachelor of Science (B.S., B.Sc. or less commonly, S.B. or Sc.B. from the Latin Scientiae Baccalaureus) is an undergraduate academic degree awarded for courses taken that generally last three years in England, Wales, Northern Ireland, Germany, India, Quebec, Singapore, Australia, New Zealand, Hong Kong and South Africa or four years in the rest of North America and Scotland. In North America, a Bachelor of Science degree usually requires students to take a majority of their courses (usually one half or three quarters) in the sciences, namely life sciences, physical sciences, or the mathematical sciences. Most universities plan the Science Bachelor degree as a liberal arts course schedule. In the UK, which subjects are considered science subjects varies, e.g. economics degrees may be B.A. in one university but B.Sc. in another. A Bachelor of Science receives the designation B.Sc. or B.S. for a major/pass degree and B.Sc. (Hons) or B.S. (Hon) for an honours degree. Formerly, in the University of Oxford the degree of B.Sc. was a postgraduate degree. This former degree, still actively granted, has since been renamed M.Sc.

"Tagged" or specified degrees

Many colleges of engineering, business, education, etc. also offer the Bachelor of Science as a tagged degree. A tagged degree incorporates the name of the major into the degree title, and generally requires more specialized course work than a degree with an untagged major. Some examples of tagged degrees include, but are not limited to:
- Bachelor of Science in Agriculture
- Bachelor of Science in Architecture
- Bachelor of Science in Business
- Bachelor of Science in Chemistry
- Bachelor of Science in Chemical Engineering
- Bachelor of Science in Civil Engineering
- Bachelor of Science in Computer Science
- Bachelor of Science in Education
- Bachelor of Science in Electrical Engineering
- Bachelor of Science in Foreign Service (BSFS)
- Bachelor of Science in Mechanical Engineering
-
Bachelor of Science in Nursing (BSN)
- Bachelor of Science in Pharmacy The Bachelor of Science in Foreign service, a degree granted by only a small number of institutions, can be found at schools (colleges) of foreign service or foreign affairs, including the
School of Foreign Service at Georgetown University, Washington, DC.

See also


-
Bachelor of Arts
-
Bachelor's degree
-
Bachelor of Mathematics
-
British undergraduate degree classification
-
British degree abbreviations Science, Bachelor

Master's degree

:For other degrees, see Academic degree or Degree (disambiguation) A master's degree is an academic degree usually awarded for completion of a postgraduate or graduate course of one to three years in duration. In the United Kingdom it is sometimes awarded for an undergraduate student whose final year consists of higher-level courses and a major research project. In the recent standardized European system of higher education diplomas, it corresponds to a two-year graduate program to be entered after three years of undergraduate studies and in preparation for either high-qualification employment or for doctoral studies.

North America

MA, MS, MSc, MSE, AM, SM

The Master of Arts (Magister Artium) and Master of Science (Magister Scientiæ) degrees are the basic type in most subjects and may be entirely course-based, entirely research-based or (more typically) a mixture. Admission to a masters program is normally contingent upon holding a bachelors degree, and progressing to a doctoral program usually requires a master's degree. In some fields or graduate programs, work on a doctorate begins immediately after the bachelors degree. Some programs provide for a joint bachelors and masters degree after about five years. Some universities use the Latin degree names, and due to the flexibility of Latin word order, Artium Magister (AM) or Scientiæ Magister (SM) may be used at some schools. For example, Harvard University uses the degree abbreviations A.M., S.M. and Ed.M. for its master's degrees.

MASc, MEng

The Master of Engineering degree is awarded to students who have done graduate work at the masters level in the field of engineering. While in the United States, candidates in engineering are typically awarded MS degrees, in the U.K. and Canada, they are generally given MSc, MASc or MEng degrees. (An example of an MEng-awarding U.S. university is the University of California, Berkeley, of which the Civil & Environmental Engineering department offers both MS and MEng degrees.) In Canada, the Master of Applied Science (MASc) is awarded to masters students with a research focus (having completed work leading to a thesis), while an MEng is awarded to masters students with a coursework focus and the completion of a research paper. The distinction between MASc and MEng is not definite since some universities grant only an MEng and some universities grant only an MASc, either research or coursework-focused.

MAT

Coursework and practica leading to a Master of Arts in Teaching (MAT) degree is intended to prepare individuals for a teaching career in a specific subject of middle and/or secondary-level curricula (i.e., middle or high school). The MAT differs from the MEd degree in that the course requirements are dominated by classes in the subject area to be taught (e.g., foreign language, math, science, etc.) rather than educational theory. Work toward most MAT degrees will, however, necessarily include classes on educational theory in order to meet program and state requirements. Work toward the MAT degree may also include practica (i.e., student teaching).

MBA, MHA, MPA, MAL

Master of Business Administration (MBA), Master of Health Administration (MHA) and Master of Public Administration (MPA), are professional degrees focusing on management for the private and public sector.

MAcc or MAc

Master of Accountancy (MAcc or MAc) is typically a one-year, non-thesis graduate program designed to prepare graduates for public accounting and to provide them with the 150 credit hours required by most states before taking the CPA exam.

MArchHist

The two-year Master of Architectural History (M.Arch.Hist.) is the first professional degree in architectural history or historic preservation and often is awarded for progress toward a Ph.D.

MDiv, MTh, STM

The Master of Divinity (M.Div.) is the first professional degree in ministry and is a common academic degree among theological seminaries. It is typically three years in length. Other theology degree titles used are Master of Theology and Master of Sacred Theology.

MEd

Master of Education degrees are similar to MA, MS, and MSc where the subject studied is education. In the United States some states license teachers with a bachelors degree but require a masters within a set number of years as continuing education. Other Education related master's degrees conferred in the United States are Master of Science in Education (M.S.Ed. or M.S.E.), Master of Arts in Education (M.A.Ed.) and Master of Adult Education (M.Ad.Ed.).

MMus

Master of Music is a three or four year degree in the field of music.

MFA

The Master of Fine Arts (MFA) is a two to three year terminal degree in a creative field of study such as theatre arts, creative writing, filmmaking or studio art.

M.A.L.S., MLA, MLS

The Master of Arts in Liberal Studies (M.A.L.S.), Master of Liberal Arts (MLA), Master of Arts in Liberal Arts (M.A.L.A.) and Master of Liberal Studies (MLS) are interdisciplinary master's degrees, occasionally awarded in specific subjects. Regardless of the title, these degrees are essentially similar, often requiring the completion of a liberal arts curriculum and a master's thesis or capstone project. In the 1950s, Wesleyan University initiated the first master's program of this sort, to refresh the educations of local secondary school teachers. Today, these degrees are often undertaken for personal enrichment, or to explore an interdisciplinary subject that does not conform to the scope of traditional master's degree programs.

MLS, MLIS, MSIS

A Master of Library Science (MLS) degree is the culmination of an interdisciplinary program encompassing information science, information management, librarianship, and/or related topics. Modern variants include Master of Library and Information Studies (MLIS), Master of Science in Information Studies (MSIS), Master of Librarianship, et al. While some universities use standard degree titles such as Master of Arts (University of Iowa) and Master of Science (University of Illinois) for their Library Science master's degrees.

MPH

The Master of Public Health degree is awarded to students who have completed a post-graduate course of study in Public Health.

MPhil

In the United States and Canada, a Master of Philosophy or Magister Philosophiae (MPhil) degree is sometimes awarded to ABD (all but dissertation) doctoral candidates who have completed all coursework, passed their written and oral examinations, and met any other special requirements before beginning work on the doctoral dissertation. Assuming all requirements are met, the MPhil degree is generally awarded after about one year of full time study towards a doctorate. The MPhil is considered equivalent to the former French DEA Diplôme d'études approfondies.

MSN, M.Nur.

The Master of Science in Nursing is the most common title for a graduate professional degree in nursing. A few schools also use the titles Master of Nursing or Master of Arts.

MSW

The Master of Social Work (MSW) is a semi-professional degree preparing students to become social workers.

 

United Kingdom

Undergraduate Masters

(MSci, MChem, MComp, MEng, MMath, MPhys, etc.) In the UK, many universities now have a four year (five years in Scotland) undergraduate programmes in science courses, with a project in the final year. The awards for these are named after the subject, so a course in mathematics would earn a Master in Mathematics degree, (abbreviated to MMath), or have a general title such as MSci (Master in Science at most universities but Master of Natural Sciences at Cambridge). Although these degrees reflect a higher level of achievement than the traditional bachelor's degree, some are generally considered less prestigious than postgraduate masters degrees such as MSc and MA. In content the first three years are generally identical to those of the equivalent bachelor's degree while the fourth year is a combination of higher-level taught courses and a research project.

Postgraduate Masters

Postgraduate masters in the United Kingdom can either be "taught" degrees, involving lectures, examination and a short dissertation, or "research" degrees (though the latter have largely been replaced by MPhil and MRes programmes, see below). Taught masters programmes involve 1 or 2 years of full-time study. The programmes are often very intensive and demanding, and concentrate on one very specialised area of knowledge. Some universities also offer a Masters by Learning Contract scheme, where a candidate can specify his or her own learning objectives; these are submitted to supervising academics for approval, and are assessed by means of written reports, practical demonstrations and presentations.

Taught Postgraduate Masters

(MSc, MA, LL.M., MLitt, MSSc, etc) The most common types of postgraduate taught Masters degrees are the Master of Arts (MA) awarded in Arts, Humanities, Theology and Social Sciences and the Master of Science (MSc) awarded in pure and applied Science. However, some universities - particularly those in Scotland - award the Master of Letters (MLitt) Master of Letters to students in the Arts, Humanities, Divinity and Social Sciences. [It should be noted that the MLitt is a research degree at the University of Cambridge, where the Master of Philosphy (MPhil) is the stanadard one-year taught degree.] In Law the standard taught degree is the Master of Laws, but certain courses may lead to the award of MA or MLitt. Until recently, both the undergraduate and postgraduate masters degrees were awarded without grade or class (like the class of an honours degree). Nowadays however, masters degrees are normally classified into the categories of Pass or Distinction, which tend to require marks of 50% and 70% respectively.

Research Postgraduate Masters

(MPhil and MRes) The Master of Philosophy (MPhil) is a research degree awarded for the completion of a thesis. It is a shorter version of the Ph.D. and some universities routinely enter potential PhD students into the MPhil programme and allow them to upgrade to the full PhD programme a year or two into the course. The Master of Research (MRes) degree is a more structured and organised version of the MPhil, usually designed to prepare a student for a career in research. For example, an MRes may combine individual research with periods of work placement in research establisments. Like the PhD, the MPhil and MRes degrees are awarded without class or grade.

MAs in Oxford, Cambridge and Dublin

The universities of Oxford, Cambridge and Dublin award masters degrees to BAs without further examination, when a certain number of years after matriculation (7 in the case of Oxford and Cambridge) have passed, and (in some but not all cases) upon payment of a nominal fee. It is commonplace for recipients of the degree to have graduated several years previously and to have had little official contact with the university or academic life since then. The only real significance of these degrees is that they historically conferred voting rights in University elections, and certain other privileges e.g. the right to dine at the holder's college's high table. They still do confer some restricted and rarely used voting rights. The MAs awarded by Oxford and Cambridge are colloquially known as the Oxbridge MA. The University of Cambridge also offers an MA to certain senior staff - both academic and non-academic - after three years' employment with the university. Until the advent of the modern research university in the mid 19th century, several other British and American universities also gave such degrees "in course".

Scottish MA

In Scotland the first degree in Arts, Fine Art, Humanities and Social Sciences awarded by many universities is the Master of Arts It should be noted the Science and Law faculties of Scottish universities award the BSc and LLB degrees respectively and the New Universities generally award the BA. The Scottish MA is roughly equivalent to a BA from a University elsewhere in the United Kingdom.

 

European Union

In order to facilitate the movement of students between European Union countries, a standardized schedule of higher education diplomas, also known as the Bologna process, was proposed: a 3-year undergraduate degree called licence or bachelors degree, then a two-year diploma called master, then a doctorate, meant to be obtained in 3 years. Because of these indicated schedules, the reform is also referred to as 3-5-3.

Austria

The traditional Austrian equivalent to the Master degree is the Diplomstudium, (At an university, only.) leading to the title Diplom-Ingenieur (female title: Diplom-Ingenieurin) in engineering or Magister (female: Magistra) in almost every other discipline. This is a first degree after 5 years of study. (The fields of Medicine, Dentistry, and Veterinary Medicine pose an exception. The first degree in these disciplines is a professional doctorate.) Due the Bologna process these first degrees are replaced by postgraduate degrees (2 years) leading to the same titles . The admission to these new degrees require a Bakkalaureus degree (female: Bakkalaura - the Austrian title for a Bachelor degree after 3 years of study) in the same or a related field or an equivalent Bachelor from other countries. The continuing use of the traditional titles for the new degrees reflects the relatively high social prestige of these titles in the Austrian society. The traditional degrees at an Austrian Fachhochschule (University of Applied Science), the Magister(FH)/Magistra(FH) and the Diplom-Ingenieur(FH)/Diplom-Ingenieurin(FH) (first degrees after 4 years of study, and not equivalent to the Master degree) are also replaced by undergraduate Bakkalaureus(FH)/Bakkalaura(FH) degrees (3 years) and postgraduate degrees (2 years, and equivalent to a Master degree) with the traditional titels. However, in some disciplines existing also postgraduate degrees with the English title Master. (e.g. the MBA in business administration or the Master of Advanced Studies, the Master of Arts, and the Master of Science in various fields of study) The admission to these new degrees also require an undergraduate degree, but not always in the same or a related discipline.

Belgium

In Belgium, owning a masters degree means that you have completed a higher education (usually university) programme of 4 or 5 years. Before the Bologna process most university degrees required 4 years of studies (leading to a licence), but some programmes required 5 years of study. An example in the field of education in business/management was the 5-year programme of "Ingénieur de Gestion" (Dutch "Handelsingenieur" - English' "Management Engineer") with an important amount of mathematics and sciences, and which corresponds to a M.Sc. in Management. This degree co-existed with an undergraduate degree in business (4 years) named "Licence en sciences économiques appliquées (Dutch. "Licentiaat in toegepaste economische wetenschappen" - English. "Licence in applied economics").

Germany

In Germany the Magister or Diplom (first degree after 5 years, from either an University or a Technische Hochschule and NOT from a Fachhochschule (University of Applied Science)) has traditionally been the equivalent to the Master degree. Due the Bologna process these academic titles are mostly being replaced by the (postgraduate) Master degree, which has caused widespread scepticism among many faculty and student body members who prefer the traditional title and curriculum. The traditional degree at a German Fachhochschule (University of Applied Science), the Diplom(FH) (first degree after 4 years of study) is also replaced by undergraduate Bachelor's degrees (3 years) and postgraduate Master's (2 years).

Finland

In Finland, the introduction of Bologna Process has standardized most of the degrees into the European model. The higher degree is called Master (of respective field) in all fields of study and takes two yeas after the Bachelor's degree. Medicine-related fields of Medicine, Dentistry, and Veterinary Medicine pose an exception. In medical fields, the licenciate (In Finnish, lisensiaatti) is an equivalent degree, the completion of which takes five (dentistry) or six years (medicine and veterinary), while the Bachelor degree is gained after second year of studies. In Engineering, the higher degree is either diplomi-insinööri (literally: "Engineer with diploma") or arkkitehti (Architect) although in international use MSc is used. In Pharmacy, the degree is proviisori. All such degrees retaining their historical name are classified as Masters. Some other Masters degrees give the right to use the traditional title of the degree-holder. E.g. the Masters of Science in Agriculture and Forestry may use the titles of metsänhoitaja or agronomi depending on their field of study.

France

In France, a traditional diploma was the maîtrise (which translates literally as "master's qualification") after 4 years of studies. This diploma becomes the first year of the Masters program, often referred to as M1. Because of this change, legal texts specifying a maîtrise (for instance, those defining the conditions for the external agrégation) had to be amended. The Masters programs subsume the former DEA (research-oriented 1-year degree), and DESS (industry-oriented 1-year degree), which become the second year of the Master (M2).

Italy

The old university system (Vecchio ordinamento) consisted in a unique five year course program, followed by a variable period (6-12 months usually) for the thesis work. This system has been reformed in 1999/2000 to adapt it to the Bologna process. Now (Nuovo Ordinamento) you have a three year course program called Laurea Breve, and after two years of specialisation, called Laurea Specialistica. Both have a final study work. Then, one can start a phd program. Medicine universities have not changed the system, and consist in six year of study followed, eventually, by the specialisation (3-6 years more).

Netherlands

In the Netherlands, the traditional acadamic degrees were doctorandus (drs.) (after 4 years; 5 years for some natural sciences, 6 years for medicine), ingenieur (ir.) (after 5 years) and for Law meester in de rechten (mr.) (after 4 years). Even though universities have adopted the masters and bachelors degree system, the old titles drs., ir. and mr. are still used (and the use of them is protected by law). The doctorandus (literally meaning "he who has to become doctor") degree is comparable with the MA degree (sometimes MSc). The ingenieur (engineer) degree is comparable with an M.Eng. or MSc degree. Finally, the mr. degree is comparable with the LL.M degree. In the Netherlands a suffix degree (MA / MSc / MEng / LL.M) can be used for holders of a prefix degree (drs., ir., mr.) instead of the prefix degree (e.g. 'ir. Jansen' or 'Jansen MSc'). A pre- and postfix can not be used at the same time (e.g. 'drs. Jansen MSc').

Poland

Currently there are two models of higher education in Poland. In the traditional model, a masters degree is awarded after completion of a university curriculum—a 5 year programme in science courses at a university or other similar institution, with a project in the final year called magisterium (it can be translated as a Master of Arts or a Master of Science thesis) that often requires carrying out research in a given field. An MA degree is called a magister (abbreviated mgr) except for medical education where it is called a lekarz medycyny (this gives the holder the right to use the title of physician) or a lekarz weterynarii in the veterinary field. Technical universities usually give the title of magister inżynier (abbreviated mgr inż.) corrseponding to an MSc Eng degree. More and more institutions introduce another model, which as of 2005 is still less popular. In this model, following the Bologna process directives, higher education is split into a 3-year bachelor programme ending with a title of licencjat (non-technical) or inżynier (technical fields), and a 2-year programme (uzupełniające studia magisterskie) giving the title of magister or magister inżynier. Nevertheless, even in these institutions, it is often possible to bridge the bachelor education directly into the master programme, without formally obtaining the licencjat degree, thus shortening the time needed for completing the education slightly. Depending on field and school, the timing may be slightly different.

Hong Kong

MA, MSc, MSocSc, MSW, MEng, LLM

Hong Kong requires two years of full-time coursework to achieve a masters degree. For part-time study, three years of study are normally required to achieve a postgraduate degree.

MPhil

As in the United Kingdom, MPhil or Master of Philosophy is a research degree awarded for the completion of a thesis, and is a shorter version of the PhD.

Taiwan

In Taiwan, bachelor degrees are basically four years (with honors). There is an entrance examination required for people who want to study in Master and PhD degrees. The course offered for Master and PhD normally is research-based.

See also


-
Bachelor's degree
-
Professional Master's degree
-
Master of Business Administration
-
Master of Engineering
-
Master of Fine Arts
-
Master of Theology
-
Licentiate
-
Engineer's degree
-
Doctorate
-
British degree abbreviations
-
Degrees of Oxford University
-
Degrees of Cambridge University

 

External links


- [http://www.ericdigests.org/pre-9210/degree.htm The Masters Degree]
- [http://www.collegeart.org/caa/ethics/mfa_standards.html College Art Association Visual Arts MFA Guidelines]

 

Normal school

A normal school is an institution for training teachers. Its purpose is to establishing teaching standards or norms, hence its name. According to the Oxford English Dictionary normal schools in the United States and Canada trained primary school teachers, while in Europe, normal schools educated primary, secondary and tertiary-level teachers. Many famous universities (including the University of California, Los Angeles) originated as normal schools. The first normal schools in the United States were founded in Massachusetts beginning in the late 1830s, thanks largely to the efforts of education reformers like Horace Mann. The first, now called Framingham State College (FSC), opened in 1839, in Lexington, Massachusetts (it later moved). Influenced by similar such academies in Prussia and elsewhere in Europe, they were intended to improve the quality of the burgeoning common school system by producing more qualified teachers. The first normal school to open west of the Appalachian Mountains in the United States was the institution now known as Illinois State University, which opened in 1857. The first to open west of the Mississippi River was the school now called Sam Houston State University, opening in 1879. The term is now archaic in many countries, and in the United States the function of normal schools has been taken up by undergraduate and graduate Schools of Education. In Canada, such institutes are usually parts of universities as the Faculty of Education offering a one- or two-year Bachelor of Education program. It requires at least three (usually four) years of prior undergraduate studies. Among countries to still use the term Normal School is New Zealand, in which normal schools are affiliated with Teachers' colleges, particularly in the cities of Christchurch and Dunedin. The term originated in the early nineteenth century from the French école normale, because the first such schools were established as standard models to be emulated by other schools. The terminology is preserved in the official translations of such schools in both the Republic of China and the People's Republic of China since the early 20th century. A Chinese normal university (, abbreviated 師大; shīdà) is usually controlled by the national or provincial government. A teachers' college (師範學院; shīfàn xuéyuàn, abbreviated 師院; shīyuàn) has lower entrance requirements. Category:Education by subject Category:History of education Category:Education in New Zealand

Race

A race is a population of humans distinguished from other populations. The most widely used racial categories are based on visible traits (especially skin color and facial features). Conceptions of race, as well as specific racial groupings, vary by culture and time and are often controversial due to their impact on social identity hence identity politics. Since the 1940s, evolutionary scientists have rejected the view of race according to which a number of finite lists of essential characteristics could be used to determine a like number of races. By the 1960s, data and models from population genetics called into question taxonomic understandings of race, and many have turned from conceptualizing and analyzing human variation in terms of race to doing so in terms of populations and clines instead. However, many scientists believe that race is a valid and useful concept. Moreover, since the 1990s, data and models from genomics and cladistics have resulted in a revolution in our understanding of human evolution, which has led some to propose a new "lineage" definition of race. These scientists have made related arguments that races are valid when understood as fuzzy sets, clusters, or extended families. Currently, opinions differ substantially within and among academic disciplines. Many evolutionary and social scientists, drawing on such biological research, think common race definitions, or any race definitions pertaining to humans, lack taxonomic rigour and validity. They argue that race definitions are imprecise, arbitrary, derived from custom, and that the races observed vary according to the culture examined. They further maintain that race is best understood as a social construct. Other scientists, however, have argued that this shift is motivated more by political than scientific reasons.

Historical origins of "race"

social construct.]]

History of the term

Given our visual acuity and complex social relationships, humans presumably have always observed and speculated about the physical differences among individuals and groups. But different societies have attributed markedly different meanings to these distinctions. The division of humanity into distinct "races" can be traced as far back as the Ancient Egyptian sacred text the Book of Gates, which identifies four categories that are now conventionally labelled "Egyptians", "Asiatics", "Libyans", and "Nubians". However, such distinctions tended to merge differences defined by features such as skin color, with tribal and national identity. Classical civilizations from Rome to China tended to invest much more importance in family or tribal affiliations than in physical appearance (Dikötter 1992; Goldenberg 2003). Ancient Greek and Roman authors also attempted to explain and categorize visible biological differences between peoples known to them. Such categories often also included fantastical human-like beings that were supposed to exist in far-away lands. Some Roman writers adhered to an environmental determinism in which climate could affect the appearance and character of groups (Isaac 2004). But in many ancient civilizations, individuals with widely varying physical appearances could become full members of a society by growing up within that society or by adopting the society's cultural norms (Snowden 1983; Lewis 1990). Medieval models of race mixed Classical ideas with the notion that humanity as a whole was descended from Shem, Ham and Japheth, the three sons of Noah, producing distinct Semitic (Asian), Hamitic (African), and Japhetic (European) peoples. The word race entered the English language in the 16th century, from French race "race, breed, lineage" (which in turn was probably a loan from Italian razza). Meanings of the term in the 16th century included "wines with a characteristic flavour", "people with common occupation", and "generation". The meaning "tribe" or "nation" emerged in the 17th century. The modern meaning, "one of the major divisions of mankind", dates to the late 18th century, but it never became exclusive (cf. continued use of "the human race"). The ultimate origin of the word is unknown; suggestions include Arabic ra'is meaning "head", but also "beginning" or "origin". The English word "race", along with many of the ideas now associated with the term, were products of the European era of exploration (Smedley 1999). As Europeans encountered people from different parts of the world, they speculated about the physical, social, and cultural differences between human groups. The rise of the African slave trade, which gradually displaced an earlier trade in slaves from throughout the world, created a further incentive to categorize human groups to justify the barbarous treatment of African slaves (Meltzer 1993). Drawing on classical sources and on their own internal interactions—for example, the hostility between the English and Irish was a powerful influence on early thinking about the differences between people (Takaki 1993)—Europeans began to sort themselves and others into groups associated with physical appearance and with deeply ingrained behaviors and capacities. A set of "folk beliefs" took hold that linked inherited physical differences between groups to inherited intellectual, behavioral, and moral qualities (Banton 1977). Although similar ideas can be found in other cultures (Lewis 1990; Dikötter 1992), they appear not to have had as much influence on social structures as they did in Europe and the parts of the world colonized by Europeans.

History of race research

The first scientific attempts to categorize race date from the 17th century, along with the development of European imperialism and colonization around the world. The first post-Classical published classification of humans into distinct races seems to be François Bernier's Nouvelle division de la terre par les différents espèces ou races qui l'habitent ("New division of Earth by the different species or races which inhabit it"), published in 1684. In the 18th century, the differences between human groups became a focus of scientific investigation (Todorov 1993). Initially, scholars focused on cataloging and describing "The Natural Varieties of Mankind," as Johann Friedrich Blumenbach entitled his 1775 text (which established the five major divisions of humans still reflected in some racial classifications). But as the science of anthropology took shape in the 19th century, European and American scientists increasingly sought explanations for the behavioral and cultural differences they attributed to groups (Stanton 1960). For example, they measured the shapes and sizes of skulls and related the results to group differences in intelligence or other attributes (Lieberman 2001). Both before and after the 1859 publication of On the Origin of Species, a debate raged in Europe over whether different human groups had the same origin or were the product of separate creations or evolutionary lineages (Wolpoff and Caspari 1997). From the 17th through the 19th centuries, the merging of folk beliefs about group differences with scientific explanations of those differences produced what one scholar has called an "ideology of race" (Smedley 1999). According to this ideology, races are primordial, natural, enduring, and distinct. Some groups might be the result of mixture between formerly distinct populations, but careful study can distinguish the ancestral races that had combined to produce admixed groups. In the 19th century a number of natural scientists wrote on race: Georges Cuvier, James Cowles Pritchard, Louis Agassiz, Charles Pickering, and Johann Friedrich Blumenbach. These scientists made three claims about race: first, that races are objective, naturally occurring divisions of humanity; second, that there is a strong relationship between biological races and other human phenomena (such as forms of activity and interpersonal relations and culture, and by extension the relative material success of cultures); third, that race is therefore a valid scientific category that can be used to explain and predict individual and group behavior. Races were distinguished by skin color, facial type, cranial profile and size, texture and color of hair. Moreover, races were almost universally considered to reflect group differences in moral character and intelligence. Their understanding of race was usually both essentialist (defining a race by a list of characteristics) and taxonomic (hierarchical). The advent of Darwinian models of evolution and Mendelian genetics, however, called into question the scientific validity of both characteristics, and required a radical reconsideration of race. The concept of race found wide application in many societies. The eugenics movement of the late 19th and early 20th centuries asserted as self-evident the biological inferiority of particular groups (Kevles 1985). In many parts of the world, the idea of race became a way of rigidly dividing groups by use of culture as well as physical appearances (Hannaford 1996). Campaigns of oppression and genocide often used supposed racial differences to motivate inhuman acts against others (Horowitz 2001).

20th- and 21st-century debates over race

Scale of race research

Discussions of race are complicated because race research has taken place on at least two scales (global and national) and from the point of view of different research aims. Evolutionary scientists are typically interested in humanity as a whole; and taxonomic racial classifications are often either unhelpful to, or refuted by, studies that focus on the question of global human diversity. Policy-makers and applied professions (such as law-enforcement or medicine), however, are typically concerned only with genetic variation at the national or sub-national scale, and find taxonomic racial categories useful. These distinctions of research aims and scale can be seen by the example of three major research papers published since 2002: Rosenberg et al. (2002), Serre & Pääbo (2004), and Tang et al. (2005). Both Rosenberg et al. and Serre & Pääbo study global genetic variation, but they arrive at different conclusions. Serre & Pääbo attribute their differing conclusions to experimental design. While Rosenberg et al. studied individuals from populations across the globe without respect to geography, Serre & Pääbo sampled individuals with respect to geography. By sampling individuals from major populations on each continent, Rosenberg et al. find evidence for genetic "clusters" (i.e., races). In contrast, Serre & Pääbo find that with respect to geography human genetic variation is continuous and "clinal". The research interest of Rosenberg et al. is medicine (i.e., epidemiology), whereas the research interest of Serre & Pääbo is human evolution. Tang et al. studied genetic variation within the United States with an interest in whether race/ethnicity or geography is of greater importance to epidemiological research. In contrast to Serre & Pääbo, Tang et al. find that race/ethnicity is of greater importance within the United States. Further [http://www.journals.uchicago.edu/AJHG/journal/issues/v77n3/42406/brief/42406.abstract.html recent research] correlating self-identified race with [http://pritch.bsd.uchicago.edu/software/structure2_1.html population genetic structure] echoed the conculsions in Tang. Indeed, the contrasting conclusions between global and national levels of analysis were predicted by Serre & Pääbo: :It is worth noting that the colonization history of the United States has resulted in a "sampling" of the human population made up largely of people from western Europe, western Africa, and Southeast Asia. Thus, studies in which individuals from Europe, sub-Saharan Africa, and Southeast Asia are used... might be an adequate description of the major components of the U.S. population.

Race as subspecies

With the advent of the modern synthesis in the early 20th century, biologists developed a new, more rigorous model of race as subspecies. For these biologists, a race is a recognizable group forming all or part of a species. A monotypic species has no races, or rather one race comprising the whole species. Monotypic species can occur in several ways:
- All members of the species are very similar and cannot be sensibly divided into biologically significant subcategories.
- The individuals vary considerably but the variation is essentially random and largely meaningless so far as genetic transmission of these variations is concerned (many plant species fit into this category, which is why horticulturists interested in preserving, say, a particular flower color avoid propagation from seed, and instead use vegetative methods like propagation from cuttings).
- The variation between individuals is noticeable and follows a pattern, but there are no clear dividing lines between separate groups: they fade imperceptibly into one another. Such
clinal variation always indicates substantial gene flow between the apparently separate groups that make up the population(s). Populations that have a steady, substantial gene flow between them are likely to represent a monotypic species even when a fair degree of genetic variation is obvious. A polytypic species has two or more races (or, in current parlance, two or more sub-types). These are separate groups that are clearly distinct from one another and do not generally interbreed (although there may be a relatively narrow hybridization zone), but which would interbreed freely if given the chance to do so. Note that groups which would not interbreed freely, even if brought together such that they had the opportunity to do so, are not races: they are separate species. Although this attempt at conceptual precision gained currency with many biologists, especially zoologists, evolutionary scientists have criticized it on a number of fronts.

The rejection of race and the rise of "population" and "cline"

At the beginning of the 20th century, anthropologists questioned, and subsequently abandoned, the claim that biologically distinct races are isomorphic with distinct linguistic, cultural, and social groups. Then, the rise of population genetics led some mainstream evolutionary scientists in anthropology and biology to question the very validity of race as scientific concept describing an objectively real phenomenon. Those who came to reject the validity of the concept, race, did so for four reasons: empirical, definitional, the availability of alternative concepts, and ethical (Lieberman and Byrne 1993). The first to challenge the concept of race on empirical grounds were anthropologists Franz Boas, who demonstrated phenotypic plasticity due to environmental factors (Boas 1912), and Ashley Montagu (1941, 1942), who relied on evidence from genetics. Zoologists Edward O. Wilson and W. Brown then challenged the concept from the perspective of general animal systematics, and further rejected the claim that "races" were equivalent to "subspecies" (Wilson and Brown 1953). One of the crucial innovations in reconceptualizing genotypic and phenotypic variation was anthropologist C. Loring Brace's observation that such variations, insofar as they are affected by natural selection, migration, or genetic drift, are distributed along geographic gradations; these gradations are called "clines" (Brace 1964). This point called attention to a problem common to phenotypic-based descriptions of races (for example, those based on hair texture and skin color): they ignore a host of other similarities and difference (for example, blood type) that do not correlate highly with the markers for race. Thus, anthropologist Frank Livingstone's conclusion that, since clines cross racial boundaries, "there are no races, only clines" (Livingstone 1962: 279). In 1964, biologists Paul Ehrlich and Holm pointed out cases where two or more clines are distributed discordantly—for example, melanin is distributed in a decreasing pattern from the equator north and south; frequencies for the haplotype for beta-S hemoglobin, on the other hand, radiate out of specific geographical points in Africa (Ehrlich and Holm 1964). As anthropologists Leonard Lieberman and Fatimah Linda Jackson observe, "Discordant patterns of heterogeneity falsify any description of a population as if it were genotypically or even phenotypically homogeneous" (Lieverman and Jackson 1995). Finally, geneticist Richard Lewontin, observing that 85 percent of human variation occurs within populations, and not between populations, argued that neither "race" nor "subspecies" was an appropriate or useful way to describe populations (Lewontin 1973). This view is purportedly debunked as Lewontin's Fallacy. Some researchers report the variation between racial groups (measured by Sewall Wright's population structure statistic FST) accounts for as little as 5% of human genetic variation2. However, because of technical limitations of FST, many geneticists now believe that low FST values do not invalidate the suggestion that there might be different human races (Edwards, 2003). Meanwhile, neo-Marxists such as David Harvey (1982, 1984, 1992) believe that race is a social construct that in reality does not exist, used instead to extenuate class differences. These empirical challenges to the concept of race forced evolutionary sciences to reconsider their definition of race. Mid-century, anthropologist William Boyd defined race as: :A population which differs significantly from other populations in regard to the frequency of one or more of the genes it possesses. It is an arbitrary matter which, and how many, gene loci we choose to consider as a significant "constellation" (Boyd 1950). Lieberman and Jackson (1994) have pointed out that "the weakness of this statement is that if one gene can distinguish races then the number of races is as numerous as the number of human couples reproducing." Moreover, anthropologist Stephen Molnar has suggested that the discordance of clines inevitably results in a multiplication of races that renders the concept itself useless (Molnar 1992). Alongside empirical and conceptual problems with "race" following the Second World War, evolutionary and social scientists were acutely aware of how beliefs about race had been used to justify discrimination, apartheid, slavery, and genocide. This questioning gained momentum in the 1960s during the U.S. civil rights movement and the emergence of numerous anti-colonial movements worldwide. In the face of these issues, some evolutionary scientists have simply abandoned the concept of race in favor of "population." What distinguishes population from previous groupings of humans by race is that it refers to a breeding population (essential to genetic calculations) and not to a biological taxon. Other evolutionary scientists have abandoned the concept of race in favor of cline (meaning, how the frequency of a trait changes along a geographic gradient). The concepts of population and cline are not, however, mutually exclusive and both are used by many evolutionary scientists. In the face of this rejection of race by evolutionary scientists, many social scientists have replaced the word race with the word "ethnicity" to refer to self-identifying groups based on beliefs in shared religion, nationality, or race. Moreover, they understood these shared beliefs to mean that religion, nationality, and race itself are social constructs and have no objective basis in the supernatural or natural realm (Gordon 1964). See also the American Anthropological Association's Statement on Race [http://www.aaanet.org/stmts/racepp.htm].

Summary of different definitions of race

The United States government has provided definitions regarding race (see for example Race (U.S. Census)). Racial classification in the U.S. 2000 census was based solely on self-identification, did not pre-suppose disjointedness, and did not include a category "Hispanic," which is considered an ethnicity, rather than a race, by the U.S. Census.

The origins, patterns, and physical manifestations of human genetic variation

Origins of modern humans

:see also single-origin hypothesis, multiregional hypothesis. multiregional hypothesis Any biological model for race must account for the development of racial differences during human evolution. For much of the 20th century, however, anthropologists relied on an incomplete fossil record for reconstructing human evolution. Their models seldom provided a firm basis for drawing inferences about the origin of races. Modern research in molecular biology, however, has provided evolutionary scientists with a whole new kind of data, which adds considerably to the knowledge of our past. There has been considerable debate among anthropologists as to the origins of Homo sapiens. About a million years ago Homo erectus migrated out of Africa and into Europe and Asia. The debate hinges on whether Homo erectus evolved into Homo sapiens more or less simultaneously in Africa, Europe, and Asia, or whether Homo sapiens evolved only in Africa, and eventually supplanted Homo erectus in Europe and Asia. Each model suggests different possible scenarios for the evolution of distinct races.

Multiregional hypothesis

Advocates of the first scenario (see Frayer et al. 1993), the multiregional continuity evolution model, cite as evidence anatomical continuity in the fossil record in South Central Europe (Smith 1982), East Asia and Australia (Wolpoff 1993) (anatomical affinity is taken to suggest genetic affinity). They argue that very strong genetic similarities among all humans do not prove recent common ancestry, but rather reflect the interconnectedness of human populations around the world, resulting in relatively constant gene flow (Thorne and Wolpoff 1992). They further argue that this model is consistent with clinal patterns (Wolpoff 1993). The most important element of this model for theories of race is that it allows a million years for the evolution of Homo sapiens around the world; this is more than enough time for the evolution of different races. Leiberman and Jackson (1995), however, have noted that this model depends on several findings relevant to race: (1) that marked morphological contrasts exist between individuals found at the center and at the perimeter of Middle Pleistocene range of the genus Homo; (2) that many features can be shown to emerge at the edge of that range before they develop at the center; and (3) that these features exhibit great tenacity through time. Regional variations in these features can thus be taken as evidence for long term differences among genus Homo individuals that prefigure different races among present-day Homo sapiens individuals.

Out of Africa

Middle Pleistocene (numbers are millennia before present).]] Information about the history of our species comes from two main sources: the paleoanthropological record and historical inferences based on current genetic differences observed in humans. Although both sources of information are fragmentary, they have been converging in recent years on the same general story. Since the 1990s, it has become common to use multilocus genotypes to distinguish different human groups and to allocate individuals to groups (Bamshad et al. 2004). These data have led to an examination of the biological validity of races as evolutionary lineages and the description of races in cladistic terms. The technique of multilocus genotyping has been used to determine patterns of human demographic history. Thus, the concept of "race" afforded by these techniques is synonymous with ancestry, broadly understood. Studies of human genetic variation imply that Africa was the ancestral source of all modern humans, and that Homo sapiens migrated out of Africa and displaced Homo erectus between 140,000 and 290,000 years ago (Cann et al. 1987). Indigenous Australians are believed to be an early out-group that remained isolated. Most other groups, including Europeans, Asians, and Native Americans, were found to be a single related (monophyletic) group resulting from a later out-migration from Africa, which could reasonably be divided into West and East Eurasian groups. The existing fossil evidence suggests that anatomically modern humans evolved in Africa, within the last ∼200,000 years, from a pre-existing population of humans (Klein 1999). Although it is not easy to define "anatomically modern" in a way that encompasses all living humans and excludes all archaic humans (Lieberman et al. 2002), the generally agreed-upon physical characteristics of anatomical modernity include a high rounded skull, facial retraction, and a light and gracile, as opposed to heavy and robust, skeleton (Lahr 1996). Early fossils with these characteristics have been found in eastern Africa and have been dated to ∼160,000–200,000 years ago (White et al. 2003; McDougall et al. 2005). At that time, the population of anatomically modern humans appears to have been small and localized (Harpending et al. 1998). Much larger populations of archaic humans lived elsewhere in the Old World, including the Neandertals in Europe and an earlier species of humans, Homo erectus, in Asia (Swisher et al. 1994). Fossils of the earliest anatomically modern humans found outside Africa are from two sites in the Middle East and date to a period of relative global warmth, ∼100,000 years ago, though this region was reinhabited by Neandertals in later millennia as the climate in the northern hemisphere again cooled (Lahr and Foley 1998). Groups of anatomically modern humans appear to have moved outside Africa permanently sometime >60,000 years ago. One of the earliest modern skeletons found outside Africa is Mungo Man, from Australia, and has been dated to ∼42,000 years ago (Bowler et al. 2003), although studies of environmental changes in Australia argue for the presence of modern humans in Australia >55,000 years ago (Miller et al. 1999). To date, the earliest anatomically modern skeleton discovered from Europe comes from the Carpathian Mountains of Romania and is dated to 34,000–36,000 years ago (Trinkaus et al. 2003). Existing data on human genetic variation support and extend conclusions based on the fossil evidence. African populations exhibit greater genetic diversity than do populations in the rest of the world, implying that humans appeared first in Africa and later colonized Eurasia and the Americas (Tishkoff and Williams 2002; Yu et al. 2002; Tishkoff and Verrelli 2003). The genetic variation seen outside Africa is generally a subset of the variation within Africa, a pattern that would be produced if the migrants from Africa were limited in number and carried just part of African genetic variability with them (Cavalli-Sforza and Feldman 2003). Patterns of genetic variation suggest an earlier population expansion in Africa followed by a subsequent expansion in non-African populations, and the dates calculated for the expansions generally coincide with the archaeological record (Jorde et al. 1998). Aspects of the relationship between anatomically modern and archaic humans remain contentious. Studies of mtDNA (Ingman et al. 2000), the Y chromosome (Underhill et al. 2000), portions of the X chromosome (Kaessmann et al. 1999), and many (though not all) autosomal regions (Harpending and Rogers 2000) support the "Out of Africa" account of human history, in which anatomically modern humans appeared first in eastern Africa and then migrated throughout Africa and into the rest of the world, with little or no interbreeding between modern humans and the archaic populations they gradually replaced (Tishkoff et al. 2000; Stringer 2002). However, several groups of researchers cite fossil and genetic evidence to argue for a more complex account. They contend that humans bearing modern traits emerged several times from Africa, over an extended period, and mixed with archaic humans in various parts of the world (Hawks et al. 2000; Eswaran 2002; Templeton 2002; Ziętkiewicz et al. 2003). As a result, they say, autosomal DNA from archaic human populations living outside Africa persists in modern populations, and modern populations in various parts of the world still bear some physical resemblance to the archaic populations that inhabited those regions (Wolpoff et al. 2001). However, distinguishing possible contributions to the gene pool of modern humans from archaic humans outside Africa is difficult, especially since many autosomal loci coalesce at times preceding the separation of archaic human populations (Pääbo 2003). In addition, studies of mtDNA from archaic and modern humans and extant Y chromosomes suggest that any surviving genetic contributions of archaic humans outside Africa must be small, if they exist at all (Krings et al. 1997; Nordborg 1998; Takahata et al. 2001; Serre et al. 2004). The observation that most genes studied to date coalesce in African populations points toward the importance of Africa as the source of most modern genetic variation, perhaps with some subdivision in the ancestral African population (Satta and Takahata 2002). Sequence data for hundreds of loci from widely distributed worldwide populations eventually may clarify the population processes associated with the appearance of anatomically modern humans (Wall 2000), as well as the amount of gene flow among modern humans since then.

Cladistics

Mungo Man A phylogenetic tree like the one shown above is usually derived from DNA or protein sequences from populations. Often mitochondrial DNA or Y chromosome sequences are used to study ancient human demographics. These single-locus sources of DNA do not recombine and are inherited from a single parent. Individuals from the various continental groups tend to be more similar to one another than to people from other continents. The tree is rooted in the common ancestor of chimpanzees and humans, which is believed to have originated in Africa. Horizontal distance corresponds to two things: #Genetic distance. Given below the diagram, the genetic difference between humans and chimps is roughly 2%, or 20 times larger than the variation among modern humans. #Temporal remoteness of the most recent common ancestor. Rough estimates are given above the diagram, in millions of years. The mitochondrial most recent common ancestor of modern humans lived roughly 200,000 years ago, latest common ancestors of humans and chimps between four and seven million years ago. Chimpanzees and humans belong to different genera, indicated in red. Formation of species and subspecies is also indicated, and the formation of "races" is indicated in the green rectangle to the right (note that only a very rough representation of human phylogeny is given). Note that vertical distances are not meaningful in this representation.

Distribution of variation

A thorough description of the differences in patterns of genetic variation between humans and other species awaits additional genetic studies of human populations and nonhuman species. But the data gathered to date suggest that human variation exhibits several distinctive characteristics. First, compared with many other mammalian species, humans are genetically less diverse—a counterintuitive finding, given our large population and worldwide distribution (Li and Sadler 1991; Kaessmann et al. 2001). For example, the chimpanzee subspecies living just in central and western Africa have higher levels of diversity than do humans (Ebersberger et al. 2002; Yu et al. 2003; Fischer et al. 2004). Two random humans are expected to differ at approximately 1 in 1000 nucleotide pairs, whereas two random chimpanzees differ at 1 in 500 nucleotide pairs. However, with a genome of approximate 3 billion nucleotides, on average two humans differ at approximately 3 million nucleotides. Most of these single nucleotide polymorphisms (SNPs) are neutral, but some are functional and influence the phenotypic differences between humans. It is estimated that about 10 million SNPs exist in human populations, where the rarer SNP allele has a frequency of at least 1% (see International HapMap Project). The distribution of variants within and among human populations also differs from that of many other species. The details of this distribution are impossible to describe succinctly because of the difficulty of defining a "population," the clinal nature of variation, and heterogeneity across the genome (Long and Kittles 2003). In general, however, 5%–15% of genetic variation occurs between large groups living on different continents, with the remaining majority of the variation occurring within such groups (Lewontin 1972; Jorde et al. 2000a; Hinds et al. 2005). This distribution of genetic variation differs from the pattern seen in many other mammalian species, for which existing data suggest greater differentiation between groups (Templeton 1998; Kittles and Weiss 2003). In the field of population genetics, it is believed that the distribution of neutral polymorphisms among contemporary humans reflects human demographic history. Our history as a species also has left genetic signals in regional populations. For example, in addition to having higher levels of genetic diversity, populations in Africa tend to have lower amounts of linkage disequilibrium than do populations outside Africa, partly because of the larger size of human populations in Africa over the course of human history and partly because the number of modern humans who left Africa to colonize the rest of the world appears to have been relatively low (Gabriel et al. 2002). In contrast, populations that have undergone dramatic size reductions or rapid expansions in the past and populations formed by the mixture of previously separate ancestral groups can have unusually high levels of linkage disequilibrium (Nordborg and Tavare 2002). In the field of population genetics, it is believed that the distribution of neutral polymorphisms among contemporary humans reflects human demographic history. It is believed that humans passed through a population bottleneck before a rapid expansion coinciding with migrations out of Africa leading to an African-Eurasian divergence around 100,000 years ago (ca. 5,000 generations), followed by a European-Asian divergence about 40,000 years ago (ca. 2,000 generations). The rapid expansion of a previously small population has two important effects on the distribution of genetic variation. First, the so-called founder effect occurs when founder populations bring only a subset of the genetic variation from their ancestral population. Second, as founders become more geographically separated, the probability that two individuals from different founder populations will mate becomes smaller. The effect of this assortative mating is to reduce gene flow between geographical groups, and to increase the genetic distance between groups. The expansion of humans from Africa affected the distribution of genetic variation in two other ways. First, smaller (founder) populations experience greater genetic drift because of increased fluctuations in neutral polymorphisms. Second, new polymorphisms that arose in one group were less likely to be transmitted to other groups as gene flow was restricted. Many other geographic, climatic, and historical factors have contributed to the patterns of human genetic variation seen in the world today. For example, population processes associated with colonization, periods of geographic isolation, socially reinforced endogamy, and natural selection all have affected allele frequencies in certain populations (Jorde et al. 2000b; Bamshad and Wooding 2003). In general, however, the recency of our common ancestry and continual gene flow among human groups have limited genetic differentiation in our species.

Substructure in the human population

genetic drift New data on human genetic variation has reignited the debate surrounding race. Most of the controversy surrounds the question of how to interpret these new data, and whether conclusions based on existing data are sound (see validity of human races). A large majority of researchers endorse the view that continental groups do not constitute different subspecies. However, other researchers still debate whether evolutionary lineages should rightly be called "races". These questions are particularly pressing for biomedicine, where self-described race is often used as an indicator of ancestry (see race in biomedicine below). Although the genetic differences among human groups are relatively small, these differences nevertheless can be used to situate many individuals within broad, geographically based groupings. For example, computer analyses of hundreds of polymorphic loci sampled in globally distributed populations have revealed the existence of genetic clustering that roughly is associated with groups that historically have occupied large continental and subcontinental regions (Rosenberg et al. 2002; Bamshad et al. 2003). Some commentators have argued that these patterns of variation provide a biological justification for the use of traditional racial categories. They argue that the continental clusterings correspond roughly with the division of human beings into sub-Saharan Africans; Europeans, western Asians, and northern Africans; eastern Asians; Polynesians and other inhabitants of Oceania; and Native Americans (Risch et al. 2002). Other observers disagree, saying that the same data undercut traditional notions of racial groups (King and Motulsky 2002; Calafell 2003; Tishkoff and Kidd 2004). They point out, for example, that major populations considered races or subgroups within races do not necessarily form their own clusters. Thus, samples taken from India and Pakistan affiliate with Europeans or eastern Asians rather than separating into a distinct cluster. However, samples from the Kalash, a small population living in northwestern Pakistan, form their own cluster on a level comparable with those of the major continental regions (Rosenberg et al. 2002). Sampling design can have a critical influence on the results of such studies. Studies of genetic clustering often have relied on samples taken from widely separated and socially defined populations. When samples were analyzed from individuals who were more evenly distributed geographically, clustering was far less evident (Serre and Pääbo 2004). Furthermore, because human genetic variation is clinal, many individuals affiliate with two or more continental groups. Thus, the genetically based "biogeographical ancestry" assigned to any given person generally will be broadly distributed and will be accompanied by sizable uncertainties (Pfaff et al. 2004). In many parts of the world, groups have mixed in such a way that many individuals have relatively recent ancestors from widely separated regions. Although genetic analyses of large numbers of loci can produce estimates of the percentage of a person's ancestors coming from various continental populations (Shriver et al. 2003; Bamshad et al. 2004), these estimates may assume a false distinctiveness of the parental populations, since human groups have exchanged mates from local to continental scales throughout history (Cavalli-Sforza et al. 1994; Hoerder 2002). Even with large numbers of markers, information for estimating admixture proportions of individuals or groups is limited, and estimates typically will have wide CIs (Pfaff et al. 2004).

Physical variation in humans

The distribution of many physical traits resembles the distribution of genetic variation within and between human populations (American Association of Physical Anthropologists 1996; Keita and Kittles 1997). For example, ∼90% of the variation in human head shapes occurs within every human group, and ∼10% separates groups, with a greater variability of head shape among individuals with recent African ancestors (Relethford 2002). A prominent exception to the common distribution of physical characteristics within and among groups is skin color. Approximately 10% of the variance in skin color occurs within groups, and ~90% occurs between groups (Relethford 2002). This distribution of skin color and its geographic patterning—with people whose ancestors lived predominantly near the equator having darker skin than those with ancestors who lived predominantly in higher latitudes—indicate that this attribute has been under strong selective pressure. Darker skin appears to be strongly selected for in equatorial regions to prevent sunburn, skin cancer, the photolysis of folate, and damage to sweat glands (Sturm et al. 2001; Rees 2003). A leading hypothesis for the selection of lighter skin in higher latitudes is that it enables the body to form greater amounts of vitamin D, which helps prevent rickets (Jablonski 2004). However, the vitamin D hypothesis is not universally accepted (Aoki 2002), and lighter skin in high latitudes may correspond simply to an absence of selection for dark skin (Harding et al. 2000). Because skin color has been under strong selective pressure, similar skin colors can result from convergent adaptation rather than from genetic relatedness. Sub-Saharan Africans, tribal populations from southern India, and Indigenous Australians have similar skin pigmentation, but genetically they are no more similar than are other widely separated groups. Furthermore, in some parts of the world in which people from different regions have mixed extensively, the connection between skin color and ancestry has been substantially weakened (Parra et al. 2004). In Brazil, for example, skin color is not closely associated with the percentage of recent African ancestors a person has, as estimated from an analysis of genetic variants differing in frequency among continent groups (Parra et al. 2003). Considerable speculation has surrounded the possible adaptive value of other physical features characteristic of groups, such as the constellation of facial features observed in many eastern and northeastern Asians (Guthrie 1996). However, any given physical characteristic generally is found in multiple groups (Lahr 1996), and demonstrating that environmental selective pressures shaped specific physical features will be difficult, since such features may have resulted from sexual selection for individuals with certain appearances or from genetic drift (Roseman 2004).

Social interpretation of physical variation

Incongruities of racial classifications

Even as the idea of "race" was becoming a powerful organizing principle in many societies, the shortcomings of the concept were apparent. In the Old World, the gradual transition in appearances from one group to adjacent groups emphasized that "one variety of mankind does so sensibly pass into the other, that you cannot mark out the limits between them," as Blumenbach observed in his writings on human variation (Marks 1995, p. 54). In parts of the Americas, the situation was somewhat different. The immigrants to the New World came largely from widely separated regions of the Old World—western and northern Europe, western Africa, and, later, eastern Asia and southern Europe. In the Americas, the immigrant populations began to mix among themselves and with the indigenous inhabitants of the continent. In the United States, for example, most people who self-identify as African American have some European ancestors—in one analysis of genetic markers that have differing frequencies between continents, European ancestry ranged from an estimated 7% for a sample of Jamaicans to ∼23% for a sample of African Americans from New Orleans (Parra et al. 1998). Similarly, many people who identify as European American have some African or Native American ancestors, either through openly interracial marriages or through the gradual inclusion of people with mixed ancestry into the majority population. In a survey of college students who self-identified as "white" in a northeastern U.S. university, ∼30% were estimated to have <90% European ancestry (Shriver et al. 2003). In the United States, social and legal conventions developed over time that forced individuals of mixed ancestry into simplified racial categories (Gossett 1997). An example is the "one-drop rule" implemented in some state laws that treated anyone with a single known African American ancestor as black (Davis 2001). The decennial censuses conducted since 1790 in the United States also created an incentive to establish racial categories and fit people into those categories (Nobles 2000). In other countries in the Americas where mixing among groups was more extensive, social categories have tended to be more numerous and fluid, with people moving into or out of categories on the basis of a combination of socioeconomic status, social class, ancestry, and appearance (Mörner 1967). Efforts to sort the increasingly mixed population of the United States into discrete categories generated many difficulties (Spickard 1992). By the standards used in past censuses, many millions of children born in the United States have belonged to a different race than have one of their biological parents. Efforts to track mixing between groups led to a proliferation of categories (such as "mulatto" and "octoroon") and "blood quantum" distinctions that became increasingly untethered from self-reported ancestry. A person's racial identity can change over time, and self-ascribed race can differ from assigned race (Kressin et al. 2003). Until the 2000 census, Latinos were required to identify with a single race despite the long history of mixing in Latin America; partly as a result of the confusion generated by the distinction, 42% of Latino respondents in the 2000 census ignored the specified racial categories and checked "some other race" (Mays et al. 2003).

Ethnicity as a way of categorizing people

As the problems surrounding the word "race" became increasingly apparent during the 20th century, the word "ethnicity" was promoted as a way of characterizing the differences between groups (Huxley and Haddon 1936; Hutchinson and Smith 1996). Ethnicity typically emphasizes the cultural, socioeconomic, religious, and political qualities of human groups rather than their genetic ancestry. It may encompass language, diet, religion, dress, customs, kinship systems, or historical or territorial identity (Cornell and Hartmann 1998). However, as a way of understanding human groups, ethnicity also suffers from several shortcomings. First, ascribing an ethnic identity to a group can imply a much greater degree of uniformity than is actually the case. In the United States, the ethnic group "Hispanic or Latino" contains such subgroups as Cuban Americans, Mexican Americans, Puerto Ricans, and recent immigrants from Central America (Hayes-Bautista and Chapa 1987). Combining these groups into a single category may serve useful bureaucratic or political ends but does not necessarily result in a better understanding of these groups. Also, ethnicity, like race, is a malleable concept that can change dramatically in different times or circumstances (Waters 1990; Smelser et al. 2001). Ethnic groups may come into existence and then dissipate as a result of broad historical or social trends. Individuals might change ethnic groups over the course of their lives or identify with more than one group. A researcher, clinician, or government official might assign an ethnicity to an individual quite different from the one that person would acknowledge (Kressin et al. 2003). Finally, despite attempts to distinguish "ethnicity" from "race," the two terms often are used interchangeably (Oppenheimer 2001). Ethnic groups can share a belief in a common ancestral origin (Cornell and Hartmann 1998), which also can be a defining characteristic of a racial group. Furthermore, ethnic groups tend to promote marriage within the group, which creates an expectation of biological cohesion regardless of whether that cohesion existed in the past.

Ancestry as a way of categorizing people

An alternative to the use of racial or ethnic categories is to categorize individuals in terms of ancestry. Ancestry may be defined geographically (e.g., Asian, sub-Saharan African, or northern European), geopolitically (e.g., Vietnamese, Zambian, or Norwegian), or culturally (e.g., Brahmin, Lemba, or Apache). The definition of ancestry may recognize a single predominant source or multiple sources. Ancestry can be ascribed to an individual by an observer, as was the case with the U.S. census prior to 1960; it can be identified by an individual from a list of possibilities or with use of terms drawn from that person's experience; or it can be calculated from genetic data by use of loci with allele frequencies that differ geographically, as described above. At least among those individuals who participate in biomedical research, genetic estimates of biogeographical ancestry generally agree with self-assessed ancestry (Tang et al. 2005), but in an unknown percentage of cases, they do not (Brodwin 2002; Kaplan 2003). race in biomedicine Genetic data can be used to infer population structure and assign individuals to groups that often correspond with their self-identified geographical ancestry. The inference of population structure from multilocus genotyping depends on the selection of a large number of informative genetic markers. These studies usually find that groups of humans living on the same continent are more similar to one another than to groups living on different continents. Many such studies are criticized for assigning group identity a priori. However, even if group identity is stripped and group identity assigned a posteriori using only genetic data, population structure can still be inferred. For example, using 377 markers, Rosenberg et al. (2002) were able to assign 1,056 individuals from 52 populations around the globe to one of six genetic clusters, of which five correspond to major geographic regions. However, in analyses that assign individuals to group it becomes less apparent that self-described racial groups are reliable indicators of ancestry. One cause of the reduced power of the assignment of individuals to groups is admixture. Some racial or ethnic groups, especially Hispanic groups, do not have homogenous ancestry. For example, self-described African Americans tend to have a mix of West African and European ancestry. Shriver et al. (2003) found that on average African Americans have ~80% African ancestry. Likewise, many white Americans have mixed European and African ancestry, where ~30% of whites have less than 90% European ancestry. In this context, it is becoming more common place to describe "race" as fractional ancestry. Without the use of genotyping, this has been approximated by the self-described ancestry of an individual's grand-parents. Nevertheless, recent research indicates that self-described race is a near-perfect indicator of an individual's genetic profile, at least in the United States. Using 326 genetic markers, Tang et al. (2005) identified 4 genetic clusters among 3,636 individuals sampled from 15 locations in the United States, and were able to correctly assign individuals to groups that correspond with their self-described race (white, African American, East Asian, or Hispanic) for all but 5 individuals (an error rate of 0.14%). They conclude that ancient ancestry, which correlates tightly with self-described race and not current residence, is the major determinant of genetic structure in the U.S. population. Genetic techniques that distinguish ancestry between continents can also be used to describe ancestry within continents. However, the study of intra-continental ancestry may require a greater number of informative markers. Populations from neighboring geographic regions typically share more recent common ancestors. As a result, allele frequencies will be correlated between these groups. This phenomenon is often seen as a cline of allele frequencies. The existence of allelic clines has been offered as evidence that individuals cannot be allocated into genetic clusters (Kittles & Weiss 2003). However, others argue that low levels of differentiation between groups merely make the assignment to groups more difficult, not impossible (Bamshad et al. 2004). Despite its seemingly objective nature, ancestry also has limitations as a way of categorizing people (Elliott and Brodwin 2002). When asked

 

 

 

 

 

 

 


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